Kenshu Shimada
Associate Professor Research Associate in Paleontology
Environmental Science Program and Sternberg Museum of Natural History
Department of Biological Sciences and Fort Hays State University
DePaul University 3000 Sternberg Drive
2325 North Clifton Avenue Hays, Kansas 67601, USA
Chicago, Illinois 60614, USA
Telephone: (773)325-4697 Fax: (773)325-7448 E-mail: kshimada@depaul.edu
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RESEARCH INTERESTS
I am a vertebrate paleontologist, a profession that deals with the anatomy and evolution of “back-boned” animals. With a background in geology (B.S. and M.S., Fort Hays State University, Hays, Kansas) and biology (Ph.D., University of Illinois at Chicago), my expertise is in fossil sharks and their paleoecology and biostratigraphy, particularly those that lived during the Cretaceous Period (i.e., the last part of “the age of dinosaurs”). Most of my research projects fall into one of three main themes. The first theme is to analyze the skeletal and dental anatomy of fossil sharks to restore their body size and body form, to make various paleobiological inferences (e.g., feeding strategies and locomotion), and to elucidate their phylogenetic relationships. My second theme is to decipher the paleoecological dynamics of extinct sharks by documenting direct or compelling evidence suggesting feeding activities and behaviors of extinct sharks (e.g., bite marks) as well as by describing co-occurring taxa, such as bony fishes and marine reptiles. My third theme is to promote paleontological applications of anatomical data taken from modern sharks (e.g., examining the quantitative relationships between the tooth size and body length in modern sharks to estimate the size of extinct sharks from teeth; examining the quality of morphological data to elucidate shark phylogeny).
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BIBLIOGRAPHY
Peer-Reviewed Articles
In press
Shimada, K., B. A. Schumacher, J. A. Parkin, and J. M. Palermo. In press. Fossil marine vertebrates from the lowermost Greenhorn Limestone (Upper Cretaceous: Middle Cenomanian) in southeastern Colorado. Journal of Paleontology Memoir.
Shimada, K., and G. L. Bell, Jr. In press. Coniasaurus Owen, 1850 (Reptilia: Squamata), from the Upper Cretaceous Niobrara Chalk of western Kansas. Journal of Paleontology, 80.
Shimada, K. In press. Types of tooth sets in the fossil record of sharks, and comments on reconstructing dentitions of extinct sharks. Journal of Fossil Research, 38(2).
Shimada, K., and M. V. Fernandes. In press. Ichthyornis sp. (Aves: Ichthyornithiformes) from the lower Turonian (Upper Cretaceous) of SEQ CHAPTER \h \r 1western Kansas. Transactions of Kansas Academy of Science.
2005
Liggett, G. A., K. Shimada, C. S. Bennett, and B. A. Schumacher. 2005. Cenomanian (Late Cretaceous) reptiles from northwestern, Russell County, Kansas. Paleobios, 25(2):9-17.
Shimada, K., and D. J. Cicimurri. 2005. Skeletal anatomy of the Late Cretaceous shark, Squalicorax (Neoselachii: Anacoracidae). Palaeontologische Zeitschrift, 79(2):241-261.
Shimada, K. 2005. Phylogeny of lamniform sharks (Chondrichthyes: Elasmobranchii) and the contribution of dental characters to lamniform systematics. Paleontological Research, 9(1):55-72.
Shimada, K., and J. F. Seigel. 2005. The relationship between the tooth size and total body length in the goblin shark, Mitsukurina owstoni (Lamniformes: Mitsukurinidae). Journal of Fossil Research, 38(1):49-56.
Blanco-Piñón, A., K. Shimada, and G. González-Barba. 2005. Lamnoid vertebrae from the Agua Nueva Formation (Upper Cretaceous: Lower-Middle Turonian), NE Mexico. Revista Mexicana de Ciencias Geológicas, 22(1):19-23.
Shimada, K. 2005 (date of imprint 2004). The relationship between the tooth size and total body length in the sandtiger shark, Carcharias taurus (Lamniformes: Odontaspididae). Journal of Fossil Research, 37(2):76-81.
2004
Shimada, K., and G. E. Hooks, III. 2004. Shark-bitten protostegid turtles from the Upper Cretaceous Mooreville Formation of Alabama. Journal of Paleontology, 78(1):205-210.
Shimada, K., K. Ewell, and M. J. Everhart. 2004. The first record of the lamniform shark genus, Johnlongia, from the Niobrara Chalk (Upper Cretaceous), western Kansas. Transactions of Kansas Academy of Science, 107(3/4):131-135.
Shimada, K., and M. J. Everhart. 2004. Shark-bitten Xiphactinus audax (Teleostei: Ichthyodectiformes) from the Niobrara Chalk (Upper Cretaceous) of Kansas. The Mosasaur, 7:35-39.
Hamm, S. A., and K. Shimada. 2004. A Late Cretaceous shark, Ptychodus martini, from Texas. Texas Journal of Science, 56(3):215-222.
2003
Shimada, K., and M. J. Everhart. 2003. Ptychodus mammillaris (Elasmobranchii) and Enchodus cf. E. shumardi (Teleostei) from the Fort Hays Limestone Member of the Niobrara Chalk (Upper Cretaceous) in Ellis County, Kansas. Transactions of Kansas Academy of Science, 106(3-4):171-176.
Shimada, K. 2003 (date of imprint 2002). The relationship between the tooth size and total body length in the white shark, Carcharodon carcharias (Lamniformes: Lamnidae). Journal of Fossil Research, 35(2):28-33.
Shimada, K., and B. A. Schumacher. 2003. The oldest record of the Late Cretaceous fish genus Thryptodus (Teleostei: Tselfatiiformes) from central Kansas. Transactions of Kansas Academy of Science. Transactions of Kansas Academy of Science, 106(1-2):54-58.
2002
Shimada, K. 2002. Dental homologies in lamniform sharks (Chondrichthyes: Elasmobranchii). Journal of Morphology, 251:38-72.
Shimada, K. 2002. Teeth of embryos in lamniform sharks (Chondrichthyes: Elasmobranchii). Environmental Biology of Fishes, 63:309-319.
Hamm, S. A., and K. Shimada. 2002. The first associated tooth set of the Late Cretaceous lamniform shark, Scapanorhynchus raphiodon (Mitsukurinidae) from the Niobrara Chalk of western Kansas. Transactions of Kansas Academy of Science, 105(1-2):18-26.
Shimada, K. 2002. Dentition of the modern basking shark, Cetorhinus maximus (Lamniformes: Cetorhinidae), and its paleontological and evolutionary implications. Journal of Fossil Research, 35(1):1-5.
Shimada, K. 2002. The relationship between the tooth size and total body length in the shortfin mako, Isurus oxyrinchus (Lamniformes: Lamnidae). Journal of Fossil Research, 35:6-9.
2001
Shimada, K. 2001. Notes on the dentition of the bigeye sandtiger shark, Odontaspis noronhai (Lamniformes: Odontaspididae). Journal of Fossil Research, 34(1):15-17.
Shimada, K. 2001. On the concept of heterodonty. Journal of Fossil Research, 34(2):52-54.
Shimada, K., and G. Hubbell. 2001. Identity of small symmetrical teeth of the Late Cretaceous lamniform shark, Cretoxyrhina mantelli, from western Kansas, U.S.A. Journal of Fossil Research, 34(2):55-57.
1999
Fielitz, C., and K. Shimada. 1999. A new species of Bananogmius (Teleostei: Tselfatiformes) from the Upper Cretaceous Carlile Shale of western Kansas. Journal of Paleontology, 73(3):504-511.
1997
Shimada, K. 1997. Periodic marker bands in vertebral centra of the Late Cretaceous lamniform shark, Cretoxyrhina mantelli. Copeia, 1997(1):233-235.
Shimada, K. 1997. Gigantic lamnoid shark vertebra from the Lower Cretaceous Kiowa Shale of Kansas. Journal of Paleontology, 71(3):522-524.
Shimada, K. 1997. Dentition of the Late Cretaceous lamniform shark, Cretoxyrhina mantelli, from the Niobrara Chalk of Kansas. Journal of Vertebrate Paleontology, 17(2):269-279.
Shimada, K. 1997. Paleoecological relationships of the Late Cretaceous lamniform shark, Cretoxyrhina mantelli (Agassiz). Journal of Paleontology, 71(5):926-933.
Shimada, K. 1997. Shark-tooth-bearing coprolite from the Carlile Shale (Upper Cretaceous), Ellis County, Kansas. Transactions of Kansas Academy of Science, 100(3-4):133-138.
Shimada, K. 1997. Stratigraphic record of the Late Cretaceous lamniform shark, Cretoxyrhina mantelli (Agassiz), in Kansas. Transactions of Kansas Academy of Science, 100(3-4):139-149.
Shimada, K. 1997. Skeletal anatomy of the Late Cretaceous lamniform shark, Cretoxyrhina mantelli, from the Niobrara Chalk in Kansas. Journal of Vertebrate Paleontology, 17(4):642-652.
1996
Shimada, K. 1996. Selachians from the Fort Hays Limestone Member of the Niobrara Chalk (Upper Cretaceous), Ellis County, Kansas. Transactions of Kansas Academy of Science, 99(1-2):1-15.
Shimada, K. 1996. Ichthyosaur (Reptilia: Ichthyosauria) vertebra from the Kiowa Shale (Lower Cretaceous: Upper Albian), Clark County, Kansas. Transactions of Kansas Academy of Science, 99(1-2):39-44.
1994
Shimada, K., and N. Inuzuka. 1994. Desmostylian tooth remains from the Miocene Tokigawa Group at Kuzubukuro, Saitama, Japan. Transactions and Proceedings of Palaeontological Society of Japan, N.S., 175:553-577.
1988
Sakamoto, O., M. Machida, T. Homma, T. Inoyama, S. Honna, and K. Shimada. 1988. Occurrence of the skeleton of Stegodon aurorae Matsumoto from Sasai, Sayama City, Central Japan. Bulletin of Saitama Museum of Natural History 6:33-44. [in Japanese with English abstract]
Abstracts
Shimada, K. 2005. Skeletal and dental anatomy of the Late Cretaceous lamniform shark, Cretalamna (=Cretolamna) appendiculata. Journal of Vertebrate Paleontology, 25(Supplement to Number 3):114A-115A.
Fernandes, M. V., and K. Shimada. 2005. SEQ CHAPTER \h \r 1A Turonian (Late Cretaceous) Bird Bone from Kansas. Abstract Kansas Academy of Science, Overland Park, [23]:18-19.
Beeson, E. R., and K. Shimada. 2004. Vertebrates from a unique bonebed of the Upper Cretaceous Niobrara Chalk, western Kansas. Journal of Vertebrate Paleontology, 24(Supplement to Number 3):37A.
Shimada, K., S. L. Cumbaa, and D. Van Rooyen. 2004. Caudal fin of the Late Cretaceous shark, Cretoxyrhina mantelli (Lamniformes: Cretoxyrhinidae). Journal of Vertebrate Paleontology, 24(Supplement to Number 3):112A.
Everhart, M. J., K. Shimada, and K. Ewell. 2004. The first record of the lamniform shark genus, Johnlongia, from the Niobrara Chalk (Upper Cretaceous), western Kansas. Abstract Missouri and Kansas Academies of Science, p. 48.
Shimada, K., J. A. Parkin, J. M. Palermo, and B. A. Schumacher. 2003. Late Cretaceous marine vertebrates from the basal Greenhorn Limestone in southeastern Colorado. Journal of Vertebrate Paleontology, 23(Supplement to Number 3):96A.
Corrado, C. A., D. A. Wilhelm, K. Shimada, and M. J. Everhart. 2003. A new skeleton of the Late Cretaceous lamniform shark, Cretoxyrhina mantelli from western Kansas. Journal of Vertebrate Paleontology, 23(Supplement to Number 3):43A.
Everhart, M. J., T. Caggiano, and K. Shimada. 2003. Note on the occurrence of five species of ptychodontid sharks from a single locality in the Smoky Hill Chalk (Late Cretaceous) of western Kansas. Abstract Kansas Academy of Science, Pittsburg, 22:[29].
Hamm, S. A., K. Shimada, and M. J. Everhart. 2003. Three uncommon lamniform sharks from the Smoky Hill Chalk (Upper Cretaceous) of western Kansas. Abstract Kansas Academy of Science, Pittsburg, 22:[30-31].
Parkin, J. A., J. M. Palermo, K. Shimada, and B. A. Schumacher. 2002. Fossil fishes from the lowermost Greenhorn Limestone (Upper Cretaceous: Middle Cenomanian) in southeastern Colorado. Geological Society of America Abstracts with Programs. [Online abstract: http://gsa.confex.com/gsa/2002AM/finalprogram/abstract_41138.htm]
Palermo, J. M., J. A. Parkin, K. Shimada, and B. A. Schumacher. 2002. Prospecting fossils from a distance: a strategy for paleontological resource management and education. Geological Society of America Abstracts with Programs. [Online abstract: http://gsa.confex.com/gsa/2002AM/finalprogram/abstract_41206.htm]
Shimada, K., M. J. Everhart, and G. E. Hooks, III. 2002. Ichthyodectid fish and protostegid turtle bitten by the Late Cretaceous lamniform shark, Cretoxyrhina mantelli. Journal of Vertebrate Paleontology, 22(Supplement to Number 3):106A.
Hanks, D. H., and K. Shimada. 2002. Vertebrate fossils, including non-avian dinosaur remains and the first shark-bitten bird bone from a Late Cretaceous (Turonian) marine deposit of northeastern South Dakota. Journal of Vertebrate Paleontology, 22(Supplement to Number 3):62A.
Schumacher, B. A., and K. Shimada. 2002. A new specimen of the Late Cretaceous teleostean fish, Thryptodus, from central Kansas, and its stratigraphic significance. Abstract Kansas Academy of Science, Hays, 21:47.
Hamm, S. A., and K. Shimada. 2002. The first associated tooth set of the extinct goblin shark, Scapanorhynchus raphiodon, from the Niobrara Chalk of western Kansas. Abstract Kansas Academy of Science, Hays, 21:32.
Shimada, K., and D. J. Cicimurri. 2001. The total length of the Late Cretaceous shark, Squalicorax (Anacoracidae). Journal of Vertebrate Paleontology, 21(Supplement to Number 3):101A.
Shimada, K. 2000. Skeletal anatomy of the Late Cretaceous shark, Squalicorax falcatus. Journal of Vertebrate Paleontology, 20(Supplement to Number 3):69A.
Shimada, K. 1999. Contribution of dental characters to elucidate the phylogeny of lamniform sharks. Journal of Vertebrate Paleontology, 19(Supplement to Number 3):75A-76A.
Shimada, K. 1998. Dental homology in macrophagous lamniform sharks. Journal of Vertebrate Paleontology, 18(Supplement to Number 3):77A.
Shimada, K. 1997. Preservation of tooth sets in the fossil record of sharks. Journal of Vertebrate Paleontology, 17(Supplement to Number 3):75A-76A.
Liggett, G. A., S. C. Bennett, K. Shimada, and J. Huenergarde. 1997. A Late Cretaceous (Cenomanian) fauna in Russell County, Kansas. Abstract Kansas Academy of Science, 16:26.
Shimada, K. 1996. Gigantic lamniform shark vertebra from the Kiowa Shale (Lower Cretaceous) of Kansas. Journal of Vertebrate Paleontology, 16(Supplement to Number 3):65A.
Everhart, M. J., P. A. Everhart, and K. Shimada. 1995. A new specimen of shark bitten mosasaur vertebrae from the Smoky Hill Chalk (Upper Cretaceous) in western Kansas. Abstract Kansas Academy of Science, Pittsburg, 14:19.
Fishman, S. E., K. Shimada, and C. Fielitz. 1995. Stratigraphic record of the Late Cretaceous genus Enchodus (Osteichthyes: Teleostei), in Kansas. Abstract Kansas Academy of Science, Pittsburg, 14:21.
Shimada, K. 1995. Skeletal anatomy of the Late Cretaceous shark, Cretoxyrhina mantelli. Journal of Vertebrate Paleontology, 15(Supplement to Number 3):53A.
Fishman, S. E., C. Fielitz, and K. Shimada. 1995. Stratigraphic record of the Late Cretaceous genus Enchodus (Osteichthyes: Teleostei), in Kansas. Journal of Vertebrate Paleontology, 15(Supplement to Number 3):28A.
Shimada, K. 1994. Scales of the Late Cretaceous shark, Cretoxyrhina mantelli, from western Kansas. Journal of Vertebrate Paleontology, 14(Supplement to Number 3):46A.
Shimada, K. 1994. Functional morphology of the dentition of the Late Cretaceous shark, Cretoxyrhina mantelli. Abstract Kansas Academy of Science, Topeka, 13:35.
Shimada, K. 1994. Occurrence of Ptychodus whipplei in Kansas and its potential diet. Abstract Kansas Academy of Science, Topeka, 13:35.
Shimada, K. 1994. Jaws of the Late Cretaceous shark, Squalicorax kaupi, from western Kansas. Journal of Morphology, 220(3):393.
Shimada, K. 1993. Upper Cretaceous elasmobranchs from the Blue Hill Shale Member of the Carlile Shale, western Kansas. Abstract Missouri and Kansas Academies of Science, Kansas City, 13:78.
Shimada, K. 1993. Upper Cretaceous selachians from the Fort Hays Limestone of the Niobrara Formation in Ellis County, Kansas. Abstract Missouri and Kansas Academies of Science, Kansas City, 13:78.
Shimada, K., and D. J. Martin. 1993. Upper Cretaceous selachians from the basal Greenhorn Limestone in Russell County, Kansas. Abstract Missouri and Kansas Academies of Science, Kansas City, 13:78.
Shimada, K. 1993. Preliminary report on the Upper Cretaceous shark, Cretoxyrhina mantelli, from western Kansas. Journal of Vertebrate Paleontology, 13(Suppl. to Number 3):57A.
Shimada, K. 1993. Jaws of the Upper Cretaceous shark, Cretoxyrhina mantelli, from western Kansas. Geological Society of America Abstracts with Programs, 25(6):58.
Theses, Technical Reports, and Miscellaneous Publications
Shimada, K., J. A. Parkin, J. M. Palermo, and B. A. Schumacher. 2003. Late Cretaceous marine vertebrates from the basal Greenhorn Limestone in southeastern Colorado. Final Project Report (unpublished) submitted to the USDA Forest Service.
Shimada, K. 1999. Dentitions of lamniform sharks: homology, phylogeny, and paleontology. Unpublished Ph.D. thesis, University of Illinois at Chicago, 486 pp.
Shimada, K. 1999. Kenshu Shimada [autobiography]. Society of Vertebrate Paleontology, News Bulletin, (176):28-29.
Shimada, K. 1994. Paleobiology of the Late Cretaceous shark, Cretoxyrhina mantelli (Lamniformes: Cretoxyrhinidae), from Kansas. Unpublished M.S. thesis, Fort Hays State University, Hays, Kansas, 169 pp.
Shimada, K. 1992. [On a missing Archaeopteryx specimen and commercial collecting of fossils]. The Newsletter of the Association for the Geological Collaboration (Japan) [Sokuho: ISSN 0388-5208], 462:6-7. [in Japanese]
Shimada, K. 1987. [Elasmobranchs from the Early Pliocene Naarai Formation, Choshi City, Chiba Prefecture, and from the Middle Miocene Tokigawa Group, Higashimatsuyama City, Saitama Prefecture, Japan]; pp. 354-357, Thirtieth Japanese Students Science Prize Complete Works, Japan Home Teacher Center, Tokyo. [in Japanese]
Shimada, K. 1986. [Elasmobranchs from the Early Pliocene Naarai Formation, Choshi City, Chiba Prefecture, Japan]; pp. 357-359, Twenty-ninth Japanese Students Science Prize Complete Works, Japan Home Teacher Center, Tokyo. [in Japanese]
(November 7, 2005)
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